(Fleming, 1823)
Marginal tentacles each with pronounced abaxial spur; with rudimentary peripheral tentacle bulbs. Umbrella up to 20 mm high, higher than wide, bell-shaped, with generally well-developed conical or spherical apical projection, thin lateral walls; manubrium of varying length, with broad base, flask-shaped; gonads typically horseshoe-shaped on whole surface of manubrium; radial canals with smooth or slightly jagged edges; mesenteries along about half length of manubrium; with 12-24, usually 16, long marginal tentacles and 16 or more club shaped marginal rudimentary bulbs, all bulbs with ocelli.