This module contains 19 tables that belong to chapter 23 of the main text, Brachyura larvae. These tables allow identification of South Atlantic brachyuran species in the larval stages, including first zoea and megalopa. The tables have been placed in a separate module in order to retain the original complex editing.

Below is a list of brachyuran species recorded from the South Atlantic, but excluding species that are endemic to
the SE Atlantic. Species marked with an asterisk (*) were not recorded along the Brazilian coast. This
list is based on Williams (1984), Boschi et al. (1992), Zolessi and Philippi (1995), Carmona and Conde
(1996), Martins and D'Incao (1996), Melo (1996), and recent additions (Rodrigues and Silva, 1998;
Sankarankutti and Ferreira, 2000). Species in bold are SW Atlantic species of brachyuran crabs with
known larval development and dealt with in this volume either as zoea or megalopa or both (see also
Table 14). Geographical and depth distributions are omitted from the original Table 1; see for those
details the species description field.
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FAMILY DROMIIDAE
Cryptodromiopsis antillensis (Stimpson, 1858)
Dromia erythropus (G. Edwards, 1771)
Hypoconcha arcuata Stimpson, 1858
Hypoconcha sabulosa (Linnaeus, 1763)= Hypoconcha parasitica

FAMILY HOMOLIDAE
Homola barbata (Fabricius, 1793)
Thelxiope barbata (Fabricius, 1793)

FAMILY LATREILLIIDAE
Latreillia elegans Roux, 1828
Latreillia williamsi Melo, 1990

FAMILY DORIPPIDAE
Ethusa americana A. Milne Edwards, 1880
Ethusa microphthalma Smith, 1881
Ethusa tenuipes Rathbun, 1897
Ethusina abyssicola Smith, 1884

FAMILY CALAPPIDAE
Acanthocarpus alexandri Stimpson, 1871
Calappa angusta A. Milne Edwards, 1880
Calappa gallus (Herbst, 1803)
Calappa nitida Holthuis, 1958
Calappa ocellata Holthuis, 1958
Calappa sulcata Rathbun, 1898
Cyclöes bairdii Stimpson, 1860
Hepatus gronovii Holthuis, 1959
Hepatus pudibundus (Herbst, 1785)
Hepatus scaber Holthuis, 1959
Osachila antillensis Rathbun, 1916
Osachila tuberosa Stimpson, 1871

FAMILY LEUCOSIIDAE
Callidactylus asper Stimpson, 1871
Ebalia rotundata (A. Milne Edwards, 1880)*
Ebalia stimpsoni A. Milne Edwards, 1880
Iliacantha intermedia Miers, 1886
Iliacantha liodactylus Rathbun, 1898
Iliacantha sparsa Stimpson, 1871
Iliacantha subglobosa Stimpson, 1871
Leucosia planata (Fabricius, 1793)*
Lithadia brasiliensis (von Martens, 1872)
Lithadia conica (Coelho, 1973)
Lithadia obliqua (Coelho, 1973)
Lithadia vertiginosa (Coelho, 1973)
Myropsis quinquespinosa Stimpson, 1871
Persephona crinita Rathbun, 1931
Persephona lichtensteinii Leach, 1817
Persephona mediterranea (Herbst, 1794)
Persephona punctata (Linnaeus, 1758)
Randallia laevis (Borradaile, 1916)
Speloeophorus elevatus Rathbun, 1898
Speloeophorus nodosus (Bell, 1855)

FAMILY CYCLODORIPPIDAE
Clythrocerus carinatus Coelho, 1973
Clythrocerus granulatus (Rathbun, 1898)
Clythrocerus moreirai Tavares, 1993
Cyclodorippe angulata Tavares, 1991
Cyclodorippe antennaria A. Milne Edwards, 1880
Deilocerus analogus (Coelho, 1973)
Deilocerus perpusillus (Rathbun, 1901)
Neocorycodus stimpsoni (Rathbun, 1937)

FAMILY CYMONOMIDAE
Cymonomoides guinotae (Tavares, 1991)
Cymonomus guillei Tavares, 1991
Cymonomus magnirostris Tavares, 1991
Cymonomus quadratus A. Milne Edwards, 1880

FAMILY RANINIDAE
Ranilia constricta (A. Milne Edwards, 1880)
Ranilia guinotae Melo and Campos Jr., 1994
Ranilia muricata H. Milne Edwards, 1837
Raninoides laevis (Latreille, 1825)
Symethis variolosa (Fabricius, 1793)

FAMILY MAJIDAE
Acanthonyx dissimulatus Coelho, 1991-1993
Acanthonyx scutiformis (Dana, 1851)
Aepinus septemspinosus (A. Milne Edwards, 1879)
Anasimus fugax A. Milne Edwards, 1880
Anasimus latus Rathbun, 1894
Anomalothir furcillatus (Stimpson, 1871)
Apiomithrax violaceus (A. Milne Edwards, 1868)
Arachnopsis filipes Stimpson, 1871
Batrachonotus brasiliensis Rathbun, 1894
Chorinus heros (Herbst, 1790)
Collodes armatus Rathbun, 1898
Collodes inermis A. Milne Edwards, 1878
Collodes rostratus A. Milne Edwards, 1878
Collodes trispinosus Stimpson, 1871
Epialtoides rostratus Coelho, 1972
Epialtus bituberculatus H. Milne Edwards, 1834
Epialtus brasiliensis Dana, 1852
Euprognatha acuta A. Milne Edwards, 1880
Euprognatha gracilipes A. Milne Edwards, 1878
Eurypodius latreillei Guérin, 1828
Hemus cristulipes A. Milne Edwards, 1875
Herbstia depressa Stimpson, 1860
Holoplites armata (A. Milne Edwards, 1880)
Inachoides forceps A. Milne Edwards, 1879
Leptopisa setirostris (Stimpson, 1871)
Leucippa pentagona H. Milne Edwards, 1833
Leurocyclus gracilipes (A. Milne Edwards and Bouvier, 1923)*
Leurocyclus tuberculosus (H. Milne Edwards and Lucas, 1843)
Libidoclaea granaria H. Milne Edwards and Lucas, 1843
Libinia bellicosa Oliveira, 1944
Libinia ferreirae Brito Capello, 1871
Libinia spinosa H. Milne Edwards, 1834
Macrocoeloma camptocerum (Stimpson, 1871)
Macrocoeloma concavum Miers, 1886
Macrocoeloma eutheca (Stimpson, 1871)
Macrocoeloma laevigatum (Stimpson, 1860)
Macrocoeloma septemspinosum (Stimpson, 1871)
Macrocoeloma subparallelum (Stimpson, 1860)
Macrocoeloma trispinosum (Latreille, 1825)
Metoporhaphis calcarata (Say, 1818)
Microlissa brasiliensis (Rathbun, 1923)
Microphrys antillensis Rathbun,1920
Microphrys bicornutus (Latreille, 1825)
Microphrys garthi (Lemos de Castro, 1953)
Microphrys interruptus Rathbun, 1920
Mithraculus coryphe (Herbst, 1801)
Mithraculus forceps (A. Milne Edwards, 1875)
Mithraculus sculptus (Lamarck, 1818)
Mithrax besnardi Melo, 1990
Mithrax braziliensis Rathbun, 1892
Mithrax caribbaeus Rathbun, 1920
Mithrax hemphilli Rathbun, 1892
Mithrax hispidus (Herbst, 1790)
Mithrax tortugae Rathbun, 1920
Mithrax verrucosus H. Milne Edwards, 1832
Mocosoa crebripunctata Stimpson, 1871
Nemausa acuticornis (Stimpson, 1871)
Nemausa cornutus (Saussure, 1857)
Nibilia antilocapra (Stimpson, 1871)
Notolopas brasiliensis Miers, 1886
Paradasygius tuberculatus (Lemos de Castro, 1949)
Pelia rotunda A. Milne Edwards, 1875
Picroceroides tubularis Miers, 1886
Pitho lherminieri (Schramm, 1867)
Podochela algicola (Stebbing, 1914)
Podochela brasiliensis Coelho, 1972
Podochela gracilipes Stimpson, 1871
Podochela minuscula Coelho, 1972
Podochela riisei Stimpson, 1860
Pyromaia tuberculata (Lockington, 1876)
Rochinia confusa Tavares, 1991
Rochinia gracilipes A. Milne Edwards, 1875
Stenocionops furcatus (Olivier, 1791)(= S. furcata)
Stenocionops spinimana (Rathbun, 1892)
Stenocionops spinosissima (Saussure, 1857)
Stenorhynchus seticornis (Herbst, 1788)
Taliepus dentatus (H. Milne Edwards, 1834)
Taliepus marginatus (Bell, 1835)
Teleophrys ornatus Rathbun, 1901
Teleophrys pococki Rathbun, 1924
Thoe aspera Rathbun, 1901
Tiche emarginata White, 1847
Tiche potiguara Garth, 1952

FAMILY PARTHENOPIDAE
Cryptopodia concava Stimpson, 1871
Heterocrypta granulata (Gibbes, 1850)
Heterocrypta lapidea Rathbun, 1901
Heterocrypta tommasii Rodrigues da Costa, 1959
Lambrus meridionalis Boschi, 1965*
Leiolambrus nitidus Rathbun, 1901
Mesorhoea sexspinosa Stimpson, 1871
Parthenope (Parthenope) agona (Stimpson, 1871)
Parthenope (Platylambrus) aylthoni (Righi, 1965)
Parthenope (Platylambrus) fraterculus (Stimpson, 1871)
Parthenope (Platylambrus) guerini (Brito Capello, 1871)
Parthenope (Platylambrus) pourtalesii (Stimpson, 1871)
Parthenope (Parthenope) serrata (H. Milne Edwards, 1834)
Solenolambrus brasiliensis Rodrigues da Costa, 1961
Solenolambrus typicus Stimpson, 1871
Thyrolambrus astroides Rathbun, 1894

FAMILY HYMENOSOMATIDAE
Halicarcinus planatus (Fabricius, 1775)*

FAMILY ATELECYCLIDAE
Peltarion spinulosum (White, 1843)

FAMILY BELLIIDAE
Acanthocyclus albatrossis Rathbun, 1898*
Acanthocyclus gayi Milne Edwards and Lucas, 1844*
Bellia picta H. Milne Edwards, 1848
Corystoides chilensis Lucas, 1844

FAMILY PORTUNIDAE
Arenaeus cribrarius (Lamarck, 1818)
Callinectes acutidens Rathbun, 1895
Callinectes bocourti A. Milne Edwards, 1879
Callinectes danae Smith, 1869
Callinectes exasperatus (Gerstaecker, 1856)
Callinectes larvatus Ordway, 1863
Callinectes ornatus Ordway, 1863
Callinectes sapidus Rathbun, 1896
Charybdis hellerii (A. Milne Edwards, 1867)
Coenophthalmus tridentatus A. Milne Edwards, 1879
Cronius ruber (Lamarck, 1818)
Cronius tumidulus (Stimpson, 1871)
Laleonectes vocans (A. Milne Edwards, 1878)
Ovalipes catharus (White, 1843)*
Ovalipes punctatus (De Haan, 1833)
Ovalipes trimaculatus (De Haan, 1833)
Portunus anceps (Saussure, 1858)
Portunus gibbesii (Stimpson, 1859)
Portunus ordwayi (Stimpson, 1860)
Portunus rufiremus Holthuis, 1959
Portunus spinicarpus (Stimpson, 1871)
Portunus spinimanus Latreille, 1819
Portunus ventralis (A. Milne Edwards, 1879)
Scylla serrata (Forskal, 1775)

FAMILY GERYONIDAE
Chaceon notialis Manning and Holthuis, 1989
Chaceon ramosae Manning, Tavares and Albuquerque, 1989

FAMILY XANTHIDAE
Actae acantha (H. Milne Edwards, 1834)
Allactaea lithostrota Williams, 1974
Banareia palmeri (Rathbun, 1894)
Carpilius corallinus (Herbst, 1783)
Cataleptodius floridanus (Gibbes, 1850)
Domecia acanthophora acanthophora (Desbonne and Schramm, 1867)
Edwardsium spinimanus (H. Milne Edwards, 1834)
Eriphia gonagra (Fabricius, 1781)
Euryozius sanguineus (Linnaeus, 1767)
Eurypanopeus abbreviatus (Stimpson, 1860)
Eurypanopeus depressus (Smith, 1869)*
Eurypanopeus dissimilis (Benedict and Rathbun, 1891)
Eurytium limosum (Say, 1818)
Garthiope barbadensis (Rathbun, 1921)
Garthiope spinipes (A. Milne Edwards, 1880)
Glyptoxanthus vermiculatus (Lamarck, 1818)
Hexapanopeus angustifrons (Benedict and Rathbun, 1891)
Hexapanopeus caribbaeus (Stimpson, 1871)
Hexapanopeus heblingi Rodrigues and Silva, 1998
Hexapanopeus manningi Sankarankutti and Ferreira, 2000
Hexapanopeus paulensis Rathbun, 1930
Hexapanopeus schmitti Rathbun, 1930
Leptodius floridanus (Gibbes, 1850)
Melybia thalamita Stimpson, 1871
Menippe nodifrons Stimpson, 1859
Micropanope nuttingi (Rathbun, 1898)
Micropanope pusilla A. Milne Edwards, 1880
Micropanope sculptipes Stimpson, 1871
Micropanope urinator (A. Milne Edwards, 1881)
Panopeus americanus Saussure, 1857
Panopeus austrobesus Williams, 1983
Panopeus bermudensis Benedict and Rathbun, 1981
Panopeus harttii Smith, 1869
Panopeus herbstii H. Milne Edwards, 1834
Panopeus lacustris Desbonne, 1867
Panopeus margentus Williams and Boschi, 1990*
Panopeus meridionalis Williams, 1983*
Panopeus occidentalis Saussure, 1857
Paractaea rufopunctata nodosa (Stimpson, 1860)
Pilumnoides coelhoi Guinot and Macpherson, 1987
Pilumnoides hassleri A. Milne Edwards, 1880
Pilumnus caribaeus Desbonne and Schramm, 1867
Pilumnus dasypodus Kingsley, 1879
Pilumnus diomedeae Rathbun, 1894
Pilumnus floridanus Stimpson, 1871
Pilumnus quoyi H. Milne Edwards, 1834
Pilumnus reticulatus Stimpson, 1860
Pilumnus spinosissimus Rathbun, 1898
Platypodiella spectabilis (Herbst, 1794)
Platyxanthus crenulatus A. Milne Edwards, 1879
Platyxanthus patagonicus A. Milne Edwards, 1879
Tetraxanthus bidentatus (A. Milne Edwards,1880)
Tetraxanthus rathbunae Chace, 1939
Xanthodius denticulatus (White, 1847)
Xanthodius parvulus (Fabricius, 1793)

FAMILY GONEPLACIDAE
Acidops cessacii (A. Milne Edwards, 1878)
Bathyplax typhla A. Milne Edwards, 1880
Chasmocarcinus cylindricus Rathbun, 1901
Chasmocarcinus peresi Rodrigues da Costa, 1968
Chasmocarcinus rathbuni Bouvier, 1917
Chasmocarcinus typicus Rathbun, 1898
Cycloplax pinnotheroides Guinot, 1969
Cyrtoplax spinidentata (Benedict, 1892)
Eucratopsis crassimanus (Dana, 1852)
Euryplax nitida Stimpson, 1859
Frevillea hirsuta (Borradaile, 1916)
Nanoplax xanthiformis (A. Milne Edwards, 1880)
Neopilumnoplax americana (Rathbun, 1898)
Panoplax depressa Stimpson, 1871
Pseudorhombila octodentata Rathbun, 1906
Pseudorhombila quadridentata (Latreille, 1828)
Speocarcinus carolinensis Stimpson, 1859
Speocarcinus meloi D'Incao and Gomes da Silva, 1992

FAMILY PINNOTHERIDAE
Clypeasterophilus stebbingi (Rathbun, 1918)
Dissodactylus crinitichelis Moreira, 1901 (= Dissodactylus encopei Rathbun, 1901)
Fabia insularis (Melo, 1971)
Fabia sebastianensis Rodrigues da Costa, 1970
Parapinnixa bouvieri Rathbun, 1918
Parapinnixa hendersoni Rathbun, 1918
Pinnaxodes chilensis (A. Milne Edwards, 1837)*
Pinnaxodes tomentosus Ortmann, 1894
Pinnixia brevipollex Rathbun, 1898 = Pinnixa rapax Bouvier, 1917
Pinnixa aidae Righi, 1967
Pinnixa chaetopterana Stimpson, 1860
Pinnixa cristata Rathbun, 1900
Pinnixa patagoniensis Rathbun, 1918
Pinnixa sayana Stimpson, 1860
Pinnotheres emiliai (Melo, 1971)
Pinnotheres garthi Fenucci, 1975
Tumidotheres maculatus (Say, 1818)
Zaops ostreum (Say, 1817)

FAMILY GRAPSIDAE
Aratus pisonii (H. Milne Edwards, 1837)
Armases angustipes (Dana, 1852)
Armases benedicti (Rathbun, 1897)
Armases miersii (Rathbun, 1897)
Chasmagnathus granulata Dana, 1851
Cyclograpsus integer H. Milne Edwards, 1837
Cyrtograpsus angulatus Dana, 1851
Cyrtograpsus affinis (Dana, 1851)
Cyrtograpsus altimanus Rathbun, 1914
Euchirograpsus americanus A. Milne Edwards, 1880
Euchirograpsus kingsleyi (Miers, 1885)
Geograpsus lividus (H. Milne Edwards, 1837)
Goniopsis cruentata (Latreille, 1803)
Grapsus grapsus (Linnaeus, 1758)= Grapsus adcensionis
Metasesarma rubripes (Rathbun, 1897)
Pachygrapsus corrugatus (Von Martens, 1872)
Pachygrapsus gracilis (Saussure, 1858)
Pachygrapsus transversus (Gibbes, 1850)
Percnon gibbesi (H. Milne Edwards, 1853)
Plagusia depressa (Fabricius, 1775)
Planes cyaneus Dana, 1851
Planes minutus (Linnaeus, 1758)*
Sesarma crassipes Cano, 1889
Sesarma rectum Randall, 1840

FAMILY GECARCINIDAE
Cardisoma guanhumi Latreille, 1825
Gecarcinus lagostoma H. Milne Edwards, 1835

FAMILY OCYPODIDAE
Ocypode quadrata (Fabricius, 1787)
Uca burgersi Holthuis, 1967
Uca cumulanta Crane, 1943
Uca leptodactyla Rathbun, 1898
Uca maracoani (Latreille, 1802-1803)
Uca mordax (Smith, 1870)
Uca rapax (Smith, 1870)
Uca thayeri Rathbun, 1900
Uca uruguayensis Nobili, 1901
Uca victoriana Von Hagen, 1987
Uca vocator (Herbst, 1804)
Ucides cordatus (Linnaeus, 1763)

FAMILY PALICIDAE
Palicus acutifrons (A. Milne Edwards, 1880)
Palicus affinis A. Milne Edwards and Bouvier, 1899
Palicus alternatus Rathbun, 1897
Palicus dentatus (A. Milne Edwards, 1880)
Palicus faxoni Rathbun, 1897
Palicus obesus (A. Milne Edwards, 1880)
Palicus sica (A. Milne Edwards, 1880)

FAMILY CRYPTOCHIRIDAE
Opecarcinus hypostegus (Shaw and Hopkins, 1977)
Troglocarcinus corallicola Verril, 1908
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