Pugh, 1995
Nectosac of anterior nectophore relatively small occupying less than half the height of the nectophore. Hydroecium without flaps, but with extensive central process. Somatocyst expanded, to a variable degree for the quarter below its middle. Posterior nectophore with relatively small nectosac. Somatocyst expanded in an irregular fashion. Extensive flap in the middle of the left hydroecial wing.