Maas, 1893
Manubrial pouches up to 29, elongate rectangular, hardly broader than space between them; lateral portions of peripheral canals very broad, transverse portions narrow. Umbrella up to 58 mm, rather flat; manubrium pouches increasing with age from 9 in juvenile specimens; gonads forming continuous folded band, interruptedly following edge of manubrium with its pouches; marginal lappets rectangular, with 2-3 statocysts; otoporpae very small.