Southwest Atlantic species of brachyuran crabs with known larval development and respective references.
Signs: +: with additional undescribed stages; *: incomplete description; ?: unknown. Numbers between brackets
include optional, additional stages.
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Species Number of larval stages Described by

Zoea Megalopa
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FAMILY DROMIIDAE
Cryptodromiopsis antillensis 6(7) 1 Rice and Provenzano, 1966
Dromia erythropus 5 1 Laughlin et al., 1982
Hypoconcha arcuata 3 1 Kircher, 1970
Hypoconcha sabulosa 3 1 Lang and Young, 1980

FAMILY HOMOLIDAE
Homola barbata 7 1 Rice and Provenzano, 1970

FAMILY LATREILLIIDAE
Latreillia elegans 2 1 Rice and Williamson, 1977; Rice, 1982

FAMILY DORIPPIDAE
Ethusa microphthalma 4 1 Martin and Truesdale, 1989

FAMILY CALAPPIDAE
Calappa gallus 1+ Taishaku and Konishi, 1995
Hepatus pudibundus 4(5)(6) 1 Rieger and Hebling, 1993

FAMILY LEUCOSIIDAE
Lithadia rotundata 1+ ? Fransozo and Bertini, 2002
Persephona mediterranea 4 1 Negreiros-Fransozo et al., 1989

FAMILY MAJIDAE
Acanthonyx scutiformis
(as Acanthonyx petiverii) 2 1 Hiyodo et al., 1994
Anasimus latus 2 1 Sandifer and van Engel, 1972
Apiomithrax violaceus 2 1 Sanatana et al., in press b
Epialtus bituberculatus 2 1 Negreiros-Fransozo and Fransozo, 2001
Epialtus brasiliensis 2 1 Negreiros-Fransozo and Fransozo, 1991
Eurypodius latreillei 2 1 Campodonico and Guzmán, 1972
Leucippa pentagona 2 1 Pohle and Marques, 2003
Libidoclaea granaria 2 1 Fagetti, 1969
Libinia ferreirae 2 1 Bakker et al., 1990
Libinia spinosa 2 1 Boschi and Scelzo, 1968;
Clark et al., 1998
Macrocoeloma camptocerum 2 1 Yang, 1967
Microphrys bicornutus 2 1 Hartnoll, 1964; Gore et al., 1982
Mithraculus coryphe 2 1 Scotto and Gore, 1980
Mithraculus forceps 2 1 Wilson et al., 1979
Mithrax caribbaeus 2 1 Bolaños et al., 1990
Mithrax hispidus 2 1 Fransozo and Hebling, 1982;
Santana et al., in press (a)
Mithrax verrucosus 2 1 Bolaños and Scelzo, 1981
Pitho lherminieri 2 1 Bolaños et al., 1996
Pyromaia tuberculata 2 1 Fransozo and Negreiros-Fransozo, 1997
Stenocionops furcatus coelatus
= Stenocionops furcata coelata 2 1 Laughlin et al., 1984;
Bolaños et al., 1994;
Santana et al., submitted
Stenorhynchus seticornis 2 1 Yang, 1976
Taliepus dentatus 2 1 Fagetti and Campodonico, 1971
Parthenope (Platylambrus) serrata
6 1 Yang, 1971

FAMILY HYMENOSOMATIDAE
Halicarcinus planatus 3 1 Boschi et al., 1969

FAMILY ATELECYCLIDAE
Peltarion spinulosum 4 1 Iorio, 1983

FAMILY BELLIIDAE
Acanthocyclus albatrossis 4 1 Campodonico and Gúzman, 1973
Acanthocyclus gayi 4 1 Fagetti and Campodonico, 1970
Corystoides chilensis 4 1 Boschi and Scelzo, 1970

FAMILY PORTUNIDAE
Arenaeus cribrarius 8 1 Stuck and Truesdale, 1988
Callinectes danae 1+ ? Sankarankutty et al., 1999
Callinectes sapidus 7(8) 1 Costlow and Bookhout, 1959
Charybdis hellerii 6 1* Negreiros-Fransozo, 1996;
Dineen et al., 2001
Coenophthalmus tridentatus 1+ 1* Boschi, 1981
Cronius ruber 1+ ? Fransozo et al., 2002
Cronius tumidulus 1+ ? Fransozo et al., 2002
Ovalipes catharus 8 1 Wear and Fielder, 1985
Ovalipes punctatus 6 1 Terada, 1980
Ovalipes trimaculatus 1+ 1* Boschi, 1981
Portunus spinicarpus 7 1 Bookhout and Costlow, 1974
Scylla serrata 5 1 Ong, 1964; Wear and Fielder, 1985

FAMILY XANTHIDAE
Carpilius corallinus 5 1* Laughlin et al., 1983
Cataleptodius floridanus 1 1* Ingle, 1987
Eriphia gonagra 4 1 Fransozo, 1987
Eurypanopeus abbreviatus 4 1 Negreiros-Fransozo, 1986a
Eurytium limosum 4 1 Kurata et al., 1981;
Messerknecht et al., 1991
Eurypanopeus depressus 4 1 Costlow and Bookhout, 1961a
Garthiope barbadensis 3(4) 1 Gore et al., 1981
Hexapanopeus angustifrons 4 1 Costlow and Bookhout, 1966a
Hexapanopeus caribbaeus 4 1 Vieira and Rieger, in press
Hexapanopeus manningi 1+ ? Hong et al., 2001
Hexapanopeus paulensis 4 1 Fransozo et al., 1989
Hexapanopeus schmitti 4 1 Bakker et al., 1989
Menippe nodifrons 5(6) 1 Scotto, 1979
Micropanope sculptipes 4 1 Andryszak and Gore, 1981
Panopeus americanus 4 1 Negreiros-Fransozo, 1986b
Panopeus austrobesus 4 1 Montú et al., 1988
Panopeus bermudensis 4 1 Martin et al., 1985
Panopeus herbstii 4 1 Costlow and Bookhout, 1961b
Panopeus occidentalis 4 1 Ingle, 1985
Pilumnus dasypodus 4 1 Sandifer, 1974;
Bookhout and Costlow, 1979
Pilumnus reticulatus 4 1 Montú et al., 1989
Platypodiella spectabilis 1+ ? Fransozo et al., 2001
Platyxanthus crenulatus 4 1 Menú-Marque, 1970; Boschi, 1981
Platyxanthus patagonicus 4 1 Iorio and Boschi, 1986
Xanthodius denticulatus 1+ 1* Lebour, 1944
Xanthodius parvulus 1+ 1* Lebour, 1944

FAMILY PINNOTHERIDAE
Clypeasterophilus stebbingi 4 1 Marques and Pohle, 1996b
Dissodactylus crinitichelis 3 1 Pohle and Telford, 1981
Pinnaxodes chilensis 1+ 1* Gutiérrez-Martínez, 1971
Pinnixa chaetopterana 1+ ? Dowds, 1980; Sandifer, 1972
Pinnixa cristata 1+ ? Dowds, 1980
Pinnixa patagoniensis 5 1 Boschi, 1981
Pinnixa sayana 1+ ? Dowds, 1980
Tumidotheres maculatus 5 1 Costlow and Bookhout, 1966b
Zaops ostreum 4 1 Hyman, 1925; Sandifer, 1972;
Sandoz and Hopkins, 1947

FAMILY GRAPSIDAE
Aratus pisonii 4 1 Warner, 1968; Díaz and Bevilacqua, 1987
Armases angustipes 4 1 Kowalczuk, 1994; Cuesta and Anger, 2001
Chasmagnathus granulata 4(5) 1 Boschi et al., 1967
Cyclograpsus integer 5(6) 1 Gore and Scotto, 1982
Cyrtograpsus angulatus 5 1 Rieger and Vieira, 1997
Cyrtograpsus altimanus 5 1 Scelzo and de Bastida, 1979
Euchirograpsus americanus 5(6) 1 Wilson, 1980
Goniopsis cruentata 1+ 1* Fransozo et al., 1998
Geograpsus lividus 1+ ? Guerao et al., 2001
Grapsus grapsus =
Grapsus adcensionis 1+ 1* Lewis, 1960; Guerao et al., 2001
Metasesarma rubripes 5 1 Díaz & Ewald, 1968; Montú et al., 1990
Pachygrapsus gracilis 13 1* Ingle, 1987;
Brossi-Garcia and Rodrigues, 1993
Pachygrapsus transversus 11 1 Cuesta and Rodrigues, 1994;
Flores et al., 1998
Percnon gibbesi 6 1 Paula and Hartnoll, 1989
Plagusia depressa 5 ? Wilson and Gore, 1980
Planes cyaneus 1+ 1* Konishi and Minagawa, 1990
Planes minutus 2 1 Lebour, 1944
Sesarma rectum 3 1 Fransozo and Hebling, 1986

FAMILY GECARCINIDAE
Cardisoma guanhumi 5 1 Costlow and Bookhout, 1968

FAMILY OCYPODIDAE
Ocypode quadrata 5 1 Díaz and Costlow, 1972
Uca burgersi 5(6) 1 Rieger, 1998
Uca leptodactyla 1+ ? Batistelli, 2003
Uca mordax 5(6) 1 Rieger, 1992
Uca rapax 1+ ? Batistelli, 2003
Uca thayeri 5 1 Anger et al., 1990
Uca uruguayensis 5(6) 1 Rieger, 1996
Uca vocator 4(5-6) 1 Rieger, 1992; Rieger, 1999
Ucides cordatus 5(6) 1 Rodrigues and Hebling, 1989

FAMILY CRYPTOCHIRIDAE
Troglocarcinus corallicola 5 ? Scotto and Gore, 1981
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